Publications
HI-Cycles: Grad Students*, Postdocs†
Updated September 2026
Submitted / In review
Section titled “Submitted / In review”Bushinsky, S. M., Nachod, Z.*, Jutras†, M., König†, D., McClish*, S., & Addey*, C. BGC-Argo+: Global biogeochemical Argo float data with secondary quality control and derived parameters. Submitted.
Bushinsky, S. M., Arteaga, L. A., Fassbender, A. J., Hauck, J., Mazloff, M. R., Cerovecki, I., et al. (n.d.). Nonlinear Interactions of Timing and Amplitude Biases in Modeled Southern Ocean pCO2: The Roles of Dissolved Inorganic Carbon, Total Alkalinity, and Sea Surface Temperature. ESS Open Archive, 2026(0110). https://doi.org/10.22541/essoar.176804975.57370146/v1
Delaigue, L., Landschützer, P., Wimart-Rousseau, C., Arbilla, L. A., Sallée, J.-B., Bushinsky, S. M., et al. (n.d.). The Southern Ocean CO2 sink in an evolving observing system. ESS Open Archive, 2026(0427). https://doi.org/10.22541/essoar.15002449/v1
Le Chevère et al., submitted.
Du, J., Cheng, L., Ito, T., Garcia, H. E., & Wang, Z. (n.d.). Climatology and annual cycle of global ocean dissolved oxygen.
Jutras†, M., Bushinsky, S. M., Cerovečki, I., Nachod, Z. *, & Hawco, N. Widespread increase of nitrate in subsurface waters over recent decades revealed by observations. ESS Open Archive, 2026(0213). https://doi.org/10.22541/essoar.177100488.81138870/v1
Published
Section titled “Published”McClish*, Shannon, Seth Bushinsky, Nathan Briggs, Ivana Cerovečki, and Clara Douglas. “Seasonal Re-Entrainment of Respired Organic Matter Decouples Southern Ocean Surface Production and Annual Net Community Production.” AGU Advances 7, no. 5 (2026): e2025AV002267. https://doi.org/10.1029/2025AV002267.
Arteaga, L. A., Rousseaux, C. S., Cetinić, I., & Bushinsky, S. (2026). Extreme 2016 El Niño heatwave weakened carbon export and respiration in the Equatorial Pacific. Communications Earth & Environment. https://doi.org/10.1038/s43247-026-03441-y
Rustogi, P., Resplandy, L., Bushinsky, S., & Deike, L. (2026). The influence of waves and bubbles on oxygen in the ocean interior. Environmental Research Letters, 21(9), 094001. https://doi.org/10.1088/1748-9326/ae612b
Deike, L., Zhou, X., Rustogi, P., Stanley, R. H. R., Reichl, B. G., Bushinsky, S. M., & Resplandy, L. (2025). A universal wind–wave–bubble formulation for air–sea gas exchange and its impact on oxygen fluxes. Proceedings of the National Academy of Sciences, 122(38), e2419319122. https://doi.org/10.1073/pnas.2419319122
Ito, T., Garcia, H. E., Wang, Z., Cheng, L., Du, J., Roach, C. J., et al. (2025). Assessing the Observational Uncertainties of Dissolved Oxygen Climatology and Seasonal Cycle Through a Coordinated Intercomparison Project. Global Biogeochemical Cycles, 39(11), e2025GB008751. https://doi.org/10.1029/2025GB008751
Bushinsky, S. M., Z. Nachod*, A. J. Fassbender, V. M. Tamsitt, Y. Takeshita, and N. Williams (2025). Offset between profiling float and shipboard oxygen observations at depth imparts bias on float pH and derived pCO2. Global Biogeochemical Cycles 39(5). DOI: 10.1029/2024GB008185. pdf.
Jutras†, M., Bushinsky, S.M., Cerovečki, I., Briggs, N (2025). Mixing accounts for more than half of biogeochemical changes along Mode Water ventilation pathways. Geophysical Research Letters 52(7). DOI: 10.1029/2024GL113789. pdf.
Dong,Y., D.C.E.Bakker, T.G.Bell,M.Yang, P.Landschützer, J.Hauck, C.Rödenbeck, V. Kitidis, S. M. Bushinsky, and P. S. Liss (2024). Direct observational evidence of strong CO2 uptake in the Southern Ocean. Science Advances 10(30). DOI: 10.1126/sciadv.adn5781
Ito, T., H. E. Garcia, Z. Wang, S. Minobe, M. C. Long, J. Cebrian, J. Reagan, T. Boyer, C. Paver, C. Bouchard, Y. Takano, S. Bushinsky, A. Cervania, and C. A. Deutsch (2024). Underestimation of multi-decadal global O2 loss due to an optimal interpolation method. Biogeosciences 21(3), pp. 747–759. DOI: 10.5194/bg-21-747-2024
Hauck, J., L. Gregor, C. Nissen, L. Patara, M. Hague, P. Mongwe, S. Bushinsky, S. C. Doney, N. Gruber, C. Le Que ́re ́, M. Manizza, M. Mazloff, P. M. S. Monteiro, and J. Terhaar (2023). The Southern Ocean Carbon Cycle 1985–2018: Mean, Seasonal Cycle, Trends, and Storage. Global Biogeochemical Cycles 37(11), e2023GB007848. DOI: 10.1029/2023GB007848
Sauve ́, J., A. R. Gray, C. J. Prend, S. M. Bushinsky, and S. C. Riser (2023). Carbon Outgassing in the Antarctic Circumpolar Current Is Supported by Ekman Transport From the Sea Ice Zone in an Observation-Based Seasonal Mixed-Layer Budget. Journal of Geophysical Research: Oceans 128(11), e2023JC019815. DOI: 10.1029/2023JC019815
Sarmiento, J. L., K. S. Johnson, L. A. Arteaga, S. M. Bushinsky, H. M. Cullen, A. R. Gray, R. M. Hotinski, T. L. Maurer, M. R. Mazloff, S. C. Riser, J. L. Russell, O. M. Schofield, and L. D. Talley (2023). The Southern Ocean carbon and climate observations and modeling (SOCCOM) project: A review. Progress in Oceanography 219, p. 103130. DOI: 10.1016/j.pocean.2023.103130
McClish*, S. and S. M. Bushinsky (2023). Majority of Southern Ocean Sea- sonal Sea Ice Zone Bloom Net Community Production Precedes Total Ice Retreat. Geophysical Research Letters. DOI: 10.1029/2023GL103459
Rodgers, K. B., J. Schwinger, A. J. Fassbender, P. Landschu ̈tzer, R. Yamaguchi, H. Fren- zel, K. Stein, J. D. Mu ̈ller, N. Goris, S. Sharma, S. Bushinsky, T.-T.-T. Chau, M. Gehlen, M. A. Gallego, L. Gloege, L. Gregor, N. Gruber, J. Hauck, Y. Iida, M. Ishii, L. Keppler, J.-E. Kim, S. Schlunegger, J. Tjiputra, K. Toyama,ˇ`P. Vaittinada Ayar, and A. Velo (2023). Seasonal Variability of the Surface Ocean Carbon Cycle: A Synthesis. Global Biogeochemical Cycles 37(9), e2023GB007798. DOI: 10.1029/2023GB007798
Sarmiento, J.L., Johnson, K.S., Arteaga, L.A., Bushinsky, S.M., Cullen, H.M., Gray, A.R., Hotinski, R.M., Maurer, T.L., Mazloff, M.R., Riser, S.C., Russell, J.L., Schofield, O.M., Talley, L.D., The Southern Ocean Carbon and Climate Observations and Modeling (SOCCOM) project: A review, Progress in Oceanography (2023), doi: https://doi.org/10.1016/j.pocean.2023.103130
Hauck J, Nissen C, Landschützer P, Rödenbeck C, Bushinsky S, Olsen A. (2023). Sparse observations induce large biases in estimates of the global ocean CO2 sink: an ocean model subsampling experiment. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences; 381(2249):-.DOI: 10.1098/rsta.2022.0063. pdf.
Huang Y, Fassbender A, Bushinsky S. Biogenic carbon pool production maintains the Southern Ocean carbon sink. (2023). Proceedings of the National Academy of Sciences. April 26; 120(18). DOI: 10.1073/pnas.2217909120. pdf.
Bushinsky, S. M., & Cerovečki, I. (2023). Subantarctic Mode Water Biogeochemical Formation Properties and Interannual Variability. AGU Advances, 4(2), e2022AV000722. doi: 10.1029/2022AV000722. pdf.
V. Tamsitt, S. Bushinsky, Z. Li, M. du Plessis, A. Foppert, S. Gille, S. Rintoul, E. Shadwick, A. Silvano, A. Sutton, S. Swart, B. Tilbrook, and N. L. Williams. Southern Ocean [in “State of the Climate in 2020”] (2021). Bull. Am. Meteorol. Soc. 102: S317–S355. doi.org/10.1175/BAMS-D-21-0081.1. pdf.
Johnson, K.S., M.F. Bif, S.M. Bushinsky, A.J. Fassbender, and Y. Takeshita. Biogeochemical Argo [in “State of the Climate in 2019”] (2020). Bull. Amer. Meteor. Soc., 101(8), S39–S41, doi: 10.1175/BAMS-D-20-0105.1. pdf.
Bushinsky, S. M., Landschützer, P., Rödenbeck, C., Gray, A. R., Baker, D., Mazloff, M. R., … Sarmiento, J. L. (2019). Reassessing Southern Ocean air-sea CO2 flux estimates with the addition of biogeochemical float observations. Global Biogeochemical Cycles, 33. https://doi.org/10.1029/2019GB006176. pdf.
Arteaga, L. A., Pahlow, M., Bushinsky, S. M., & Sarmiento, J. L. (2019). Nutrient Controls on Export Production in the Southern Ocean. Global Biogeochemical Cycles, 33(8), 942–956. https://doi.org/10.1029/2019gb006236.
Bushinsky, S. M., Takeshita, Y., & Williams, N. L. (2019). Observing Changes in Ocean Carbonate Chemistry: Our Autonomous Future. Current Climate Change Reports. https://doi.org/10.1007/s40641-019-00129-8. pdf
Bushinsky, S. M., & Emerson, S. R. (2018). Biological and physical controls on the oxygen cycle in the Kuroshio Extension from an array of profiling floats. Deep Sea Research Part I: Oceanographic Research Papers, 141, 51–70. https://doi.org/10.1016/j.dsr.2018.09.005. pdf
Gray, A. R., Johnson, K. S., Bushinsky, S. M., Riser, S. C., Russell, J. L., Talley, L. D., … Sarmiento, J. L. (2018). Autonomous Biogeochemical Floats Detect Significant Carbon Dioxide Outgassing in the High-Latitude Southern Ocean. Geophysical Research Letters, 45(17), 9049–9057. https://doi.org/10.1029/2018GL078013. pdf. Supporting Information.
Bushinsky, S. M., Gray, A. R., Johnson, K. S., & Sarmiento, J. L. (2017). Oxygen in the Southern Ocean from Argo floats: Determination of processes driving air-sea fluxes. Journal of Geophysical Research: Oceans, 122, 1–22. https://doi.org/10.1002/2017JC012923. pdf. Supporting Information.
Yang, B., Emerson, S. R., & Bushinsky, S. M. (2017). Annual net community production in the subtropical Pacific Ocean from in-situ oxygen measurements on profiling floats. Global Biogeochemical Cycles, 31(4), 728–744. https://doi.org/10.1002/2016GB005545. pdf
Bushinsky, S. M., Emerson, S. R., Riser, S. C., & Swift, D. D. (2016). Accurate oxygen measurements on modified Argo floats using in situ air calibrations. Limnology and Oceanography: Methods, 00(609), 1–48. https://doi.org/10.1002/lom3.10107. pdf
Newsom, E. R., Fassbender, A. J., Maloney, A. E., & Bushinsky, S. M. (2016). Increasing the usability of climate science in political decision-making. Elementa: Science of the Anthropocene, 4, 000127. https://doi.org/10.12952/journal.elementa.000127. pdf
Emerson, S. R., & Bushinsky, S. (2016). The role of bubbles during air-sea gas exchange. Journal of Geophysical Research: Oceans, 121(6), 4360–4376. https://doi.org/10.1002/2016JC011744. pdf
Bushinsky, S. M., & Emerson, S. R. (2015). Marine Biological Production from In Situ Oxygen Measurements on a Profiling Float in the Subarctic Pacific Ocean. Global Biogeochemical Cycles, 29. https://doi.org/10.1002/2015GB005251. pdf
Bushinsky, S. M. (2015). Improved estimates of air-sea oxygen fluxes and biological carbon export through the use of self-calibrating Argo oxygen floats in the Pacific. University of Washington. Thesis. http://hdl.handle.net/1773/35282. pdf
Emerson, S., & Bushinsky, S. (2014). Oxygen Concentrations and Biological Fluxes in the Open Ocean. Oceanography, 27(1), 168–171. https://doi.org/10.5670/oceanog.2014.20. pdf
Bushinsky, S. M., & Emerson, S. (2013). A method for In-situ Calibration of Aanderaa Oxygen Sensors on Surface Moorings. Marine Chemistry, 155, 22–28. https://doi.org/10.1016/j.marchem.2013.05.001. pdf
Howard, E., Emerson, S., Bushinsky, S., & Stump, C. (2010). The role of net community production in air-sea carbon fluxes at the North Pacific subarctic-subtropical boundary region. Limnology and Oceanography, 55(6), 2585–2596. https://doi.org/10.4319/lo.2010.55.6.2585. pdf
Micheli, F., Shelton, A. O., Bushinsky, S. M., Chiu, A. L., Haupt, A. J., Heiman, K. W., … Watanabe, J. (2008). Persistence of depleted abalones in marine reserves of central California. Biological Conservation, 141(4), 1078–1090. https://doi.org/10.1016/j.biocon.2008.01.014. pdf
Arrigo, K. R., van Dijken, G. L., & Bushinsky, S. (2008). Primary production in the Southern Ocean, 1997-2006. Journal of Geophysical Research, 113(C08004). https://doi.org/10.1029/2007JC004551. pdf
Baglole, C. J., Bushinsky, S. M., Garcia, T. M., Kode, A., Rahman, I., Sime, P. J., & Phipps, R. P. (2006). Differential induction of apoptosis by cigarette smoke extract in primary human lung fibroblast strains: implications for emphysema. Am J Physiol Cell Mol Physiol, 291, L19–L29. https://doi.org/10.1152/ajplung.00306.2005. pdf